Insects that have displaced most gnats? (4)

I believe the answer is:

ants

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'insects that' is the definition.
'ants' can be an answer for 'insects' (ant is a kind of insect). I am not sure about the remainder of the definition.

'displaced most gnats?' is the wordplay.
'displaced' is an anagram indicator.
'most' means to remove the first letter.
'gnats' with its initial letter removed is 'nats'.
'nats' with letters rearranged gives 'ANTS'.

'have' is the link.

Can you help me to learn more?

(Other definitions for ants that I've seen before include "Soldiers" , "Tiny insects that live in colonies" , "Busy little insects" , "Small social insects" , "Industrious insects" .)

I've seen this clue in the Evening Standard.

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