By contrast, the benefits associated with feathers turned right over (6,4)
I believe the answer is:
upside down
'over' is the definition.
(thesaurus)
'by contrast the benefits associated with feathers turned right' is the wordplay.
I cannot quite see how this works, but
'by' could be 'side' (synonyms) and 'side' is found within the answer.
'with' could be 'w' (abbreviation) and 'w' is present in the answer.
This explanation may well be incorrect...
Can you help me to learn more?
(Other definitions for upside down that I've seen before include "in a chaotic manner" , "in disarray" , "in a crazy way" , "The wrong way up" , "Inverted" .)