Comfortable part of ecosystem (4)
I believe the answer is:
cosy
'comfortable' is the definition.
(similar in meaning)
'part of ecosystem' is the wordplay.
'part of' indicates a hidden word.
'COSY' is hidden in the letters of 'ecosystem'.
(Other definitions for cosy that I've seen before include "Warm, secure" , "Comfortable; teapot cover" , "Comfy and snug" , "Warm and snug" , "Warm and comfortable" .)