Express anger (5)
(Other definitions for growl that I've seen before include "Sound of hostile animal" , "threatening sound" , "Menacing animal sound" , "snap" , "Hostile animal's sound" .)
(Other definitions for fume that I've seen before include "Express great anger" , "Chafe" , "Smoke and be angry" , "Feel great anger" , "Champ at the bit" .)