National symbol of kingdom in West Indies, originally (4)
I believe the answer is:
kiwi
'national symbol' is the definition.
(the kiwi bird is a symbol of New Zealand)
'kingdom in west indies originally' is the wordplay.
'originally' indicates taking the first letters.
The initial letters of 'kingdom in west indies' is 'KIWI'.
'of' acts as a link.
(Other definitions for kiwi that I've seen before include "one with big bill" , "New Zealand bird that does not fly" , "Fruit and flightless bird" , "Symbol of New Zealand" , "Sacred Japanese peak" .)