One settling on bark after wind (5)
I believe the answer is:
payer
'one settling' is the definition.
(eg someone settling a bill)
'on bark after wind' is the wordplay.
'on' becomes 're' ('on' can be mean 'with reference to').
'bark' becomes 'yap' (yapping is a kind of barking).
'after wind' says the letters should be written in reverse (after being wound back).
're'+'yap'='reyap'
'reyap' written backwards gives 'PAYER'.
(Other definitions for payer that I've seen before include "Who provides money" , "Remitter" , "Someone to fund" , "One handing over money" , "One foots the bill" .)