Crossword Genius

Pair, say, up in pain (6)

Ross

I believe the answer is:

twinge

'pain' is the definition.
(twinge is a kind of pain)

'pair say up' is the wordplay.
'pair' becomes 'twin' ('twin' can be a synonym of 'pair'**).
'say' becomes 'eg'.
'up' is a reversal indicator (in down clue: letters go upwards).
'eg' back-to-front is 'ge'.
'twin'+'ge'='TWINGE'

'in' is the link.

(Other definitions for twinge that I've seen before include "Brief sharp pain" , "Brief sudden pain" , "Spasm" , "Sharp pang" , "Sudden stab of pain" .)

I've seen this clue in The Telegraph.
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