Prevailing feeling is low key (4)
I believe the answer is:
mood
'prevailing feeling' is the definition.
'mood' can be an answer for 'feeling' (mood is a kind of feeling). I am unsure of the 'prevailing' bit.
'low key' is the wordplay.
'low' becomes 'moo' (both can mean to make a cow noise).
'key' becomes 'd' (musical key).
'moo'+'d'='MOOD'
'is' is the link.
Can you help me to learn more?
(Other definitions for mood that I've seen before include "at times appears very high" , "Temporary state of mind" , "Humour - bad temper" , "State of mind or feeling" , "melancholy" .)