They enable us to hold our heads high (5)

I believe the answer is:

necks

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'they' is the definition.
Both the definition and answer are plural nouns.
Maybe you can see an association between them that I don't see?

I don't understand how the remainder of the clue works.

Can you help me to learn more?

(Other definitions for necks that I've seen before include "behaves amorously" , "Smooches" , "They connect heads to bodies" , "Features of swans and giraffes" , "Kisses amorously" .)

I've seen this clue in the Evening Standard.

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