Crossword Genius

Time scoundrels spread out keeping the swine on his toes (8)

Ross

I believe the answer is:

trotters

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'toes' is the definition.
Although both the answer and definition are plural nouns, I can't understand how one could define the other.

I cannot understand how the rest of the clue works.

Can you help me to learn more?

(Other definitions for trotters that I've seen before include "They carry pigs" , "Sitcom family" , "trained horses" , "Sounder" , "Racing horses or pigs feet" .)

I've seen this clue in the Irish Times (Crosaire) and the Irish Times.
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