Titular character in the best-selling DVD of all time (4)

I believe the answer is:

nemo

(Other definitions for nemo that I've seen before include "Fish sought in film" , "- me impune lacessit (motto of Scotland)" , "commander for the Nautilus" , "Finding -, animated film" , "Jules Verne's submarine captain" .)

I've seen this clue in The New York Times.

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