Understand and run in (9)
I believe the answer is:
apprehend
I believe this is a double definition.
'understand' is the first definition.
(apprehending is a kind of understanding)
'run in' is the second definition.
(thesaurus)
'and' acts as a link.
(Other definitions for apprehend that I've seen before include "Catch; understand" , "Arrest - grasp" , "Recognise - arrest - suspect" , "Fear - arrest" , "bust" .)