Where the caterpillars have been in the greens? (5)
I believe the answer is:
holes
'greens?' is the definition.
The answer and definition can be both natural objects as well as being plural nouns.
Perhaps you can see a link between them that I don't see?
I can't explain the rest of the clue.
Can you help me to learn more?
(Other definitions for holes that I've seen before include "Cavities" , "Scrapes" , "Openings" , "Apertures" , "Empty spaces" .)