They're always under your feet (5)

I believe the answer is:

soles

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'feet' is the definition.
The definition and answer can be both animals as well as being plural nouns.
Maybe there's a link between them I don't understand?

I don't understand how the rest of the clue works.

Can you help me to learn more?

(Other definitions for soles that I've seen before include "Undersides of feet" , "Shoe bottoms" , "They're always under one's feet!" , "Undersurfaces of feet" , "Parts of shoes" .)

I've seen this clue in the Evening Standard.

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