Crossword Genius

Vigilant dancing in Swan Lake? (2,4,4)

Ross

I believe the answer is:

on ones toes

'vigilant' is the definition.
(as in being kept on one's toes)

I don't understand how the remainder of the clue works.

(Other definitions for on ones toes that I've seen before include "ready" , "Alert; where the interfering tread" .)

I've seen this clue in The Guardian.
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